diff --git a/src/ngMock/angular-mocks.js b/src/ngMock/angular-mocks.js
index 3910cca15baa..6a391832c638 100644
--- a/src/ngMock/angular-mocks.js
+++ b/src/ngMock/angular-mocks.js
@@ -797,7 +797,10 @@ angular.mock.animate = angular.module('ngAnimateMock', ['ng'])
 
       var animateJsConstructor = function() {
         var animator = $delegate.apply($delegate, arguments);
-        runners.push(animator);
+        // If no javascript animation is found, animator is undefined
+        if (animator) {
+          runners.push(animator);
+        }
         return animator;
       };
 
diff --git a/test/ngMock/angular-mocksSpec.js b/test/ngMock/angular-mocksSpec.js
index 818d33e3003a..cdee629c6d40 100644
--- a/test/ngMock/angular-mocksSpec.js
+++ b/test/ngMock/angular-mocksSpec.js
@@ -2230,12 +2230,34 @@ describe('ngMockE2E', function() {
         expect(animationLog).toEqual(['start leave', 'end leave']);
       }));
 
+      it('should not throw when a regular animation has no javascript animation',
+        inject(function($animate, $$animation, $rootElement) {
+
+        var element = jqLite('<div></div>');
+        $rootElement.append(element);
+
+        // Make sure the animation has valid $animateCss options
+        $$animation(element, null, {
+          from: { background: 'red' },
+          to: { background: 'blue' },
+          duration: 1,
+          transitionStyle: '1s linear all'
+        });
+
+        expect(function() {
+          $animate.closeAndFlush();
+        }).not.toThrow();
+
+        dealoc(element);
+      }));
+
       it('should throw an error if there are no animations to close and flush',
         inject(function($animate) {
 
         expect(function() {
           $animate.closeAndFlush();
         }).toThrow('No pending animations ready to be closed or flushed');
+
       }));
     });
   });